Viral Vector Titer Methods: Capsid, Genome & Infectious Titers Explained

Two jars of marbles that are color-coded to indicate the total amount of particles in the jar (left) vs. the number of infectious particles (right, a subset of the total)

When someone asks, “How much vector do we have?” the answer is never as simple as a single number.

That’s because not all viral particles are created equal. Some are functional, delivering their genetic payload like champions. Others are just along for the ride - empty shells or defective copies that look like the part but don’t do the job.

Understanding this distinction is critical. A batch may look abundant, but what really matters is how many particles are truly infectious, and how that relates to the total number of viral particles (Particle:Infectivity ratio).

3 Ways We Measure Yield

Capsid Titer (Protein-based)

What it is: Measures how many viral “shells” (capsids) are present, regardless of what’s inside.

Example: For lentiviral vectors, a common method is the p24 ELISA, which detects a structural protein in the viral capsid.

What it tells you: “How many virus particles were made?”

Limitations: Doesn’t distinguish between full vs. empty particles, or between infectious vs. defective ones.

Schematic illustrating how genome titer assays only count viral vector particles containing payload

Genome Titer

What it is: Counts how many viral genomes are packaged.

Example: For AAV, qPCR against ITR sequence is standard.

What it tells you: “How many genomes are present inside capsids?” Since each viral particle packages a specific number of genome copies (1 for AAV, 2 for Lentiviral vector), you can use this to determine how many viral particles you have.

Limitations: Can’t tell if the packaged genome is intact, or if the virus is functional once it enters a cell.

Illustration showing how capsid ELISAs count every viral vector particle

Infectious Titer (Function-based)

What it is: Measures the fraction of particles that actually infect cells and deliver the genetic payload.

Example: For lentiviral vectors, cell-based assays with a qPCR or flow-based readout can directly assess functionality.

What it tells you: “How many particles are doing the job we care about?”

Limitations: Cell-based assays are more variable and have longer turnaround time than ELISA or PCR-based methods. In addition, the assay design must be specific to the vector system and payload.

Illustration of a viral vector potency assay to measure functional viral particles

Why They Don’t Match

Even if you measure the same batch of vector by all three methods, the numbers will differ - sometimes dramatically. Why?

  • Not every capsid contains a genome

  • Not every genome is full intact or packaged correctly

  • Not every correctly packaged viral particle can infect/transduce a cell

Think of it like a factory producing old-school light bulbs:

  • Capsid titer = How many bulbs roll off the line

  • Genome titer = How many bulbs have filaments inside

  • Infectious titer = How many bulbs can actually light up

Illustration of lightbulbs on a factory. Used as a metaphor for describing viral vector analytical strategy. conveyor belt; only some of them are functional

Why It Matters

The difference between particle counts and infectious units isn’t just technical hair-splitting. It has real-world consequences:

  • Dosing: Patients need infectious particles, not just total particles. A misleading count could mean under- or over-dosing.

  • Safety: Non-infectious particles can still trigger immune responses, complicating therapy.

  • Manufacturing Strategy: High particle-to-infectivity ratios can point to manufacturing inefficiencies that should be addressed upstreams (e.g. by varying plasmid ratios or transfection conditions).

Schematic showing that a well-designed vector analytical strategy impacts dosing, safety, and viral vector manufacturing

Key Takeaways

  • Not all viral particles are infectious

  • Different assays tell different parts of the story

  • Robust viral vector analytics are essential for accurate dosing, safety, and regulatory success

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